*1: Gooseberry bush in
orchard on Newlyn Cliff, 1863, WC (PNZ)
(Paton 1969a: 745); also: base of sallows in stream, Poltesco
valley, Lizard, June 1962, JAP (Warburg 1963b: 501, Paton
*2: By the R. Camel,
Boscarne near Bodmin, 1879, RVT (B) (Paton
The only record in West Penwith (Newlyn Cliff,
SW42U) is from 1863 and was on a Gooseberry bush in an
orchard, possibly an introduction.
Grows in mats. Commonest on silted bark in
inundation-zones of rivers or large streams, on bark (Alder,
Ash, Sycamore, Blackthorn, etc.) or sometimes rock (including
serpentinite), often where partly shaded. Also on tree bases
Elsewhere, extends outside flood-zones in some humid and
sheltered locations, occurring with other epiphytes e.g. on
sheltered branches of coppiced Hazels on open wooded slopes.
One record low in old wet Grey Willow carr (on Goss Moor).
Common associates include Kindbergia praelonga,
trichomanoides, Metzgeria furcata, Orthotrichum rivulare,
more rarely Dendrocryphaea
Commonly c.fr.: capsules immature 1, 3, 5, 7;
dehiscing 6, 7; dehisced [old 3, 5] 10.