*1: Carbis
Bay,
1886, WC & HND (GL) (Paton 1969a:
726).
*2: Pendower
Beach
near Veryan, 1961, JAP (BBSUK) (Paton 1969a:
726).
Grows as patches or more extensive low lawns.
Locally common on calcareous sand in short moss-rich dune
grassland, and where calcareous blown-sand lies on coastal
hillslopes, above sea-cliffs, or on old mine-spoil. Also
locally and in small amounts in low vegetation of open slopes
above or near rocky sea-cliffs (serpentinite, slates) and in a
slate quarry. Restricted in Cornwall to coasts or sites
on dunes within 1 km inland of coasts. Normally in fully
insolated places (occasionally part-shaded where colonising
bushes encroach onto its duneland habitats). Associates on
dunes and sandy coastal slopes include Barbula convoluta, Bryum spp., Ditrichum gracile, Syntrichia ruralis
var.
ruraliformis, Tortella flavovirens,
Trichostomum
brachydontium, Trichostomum
crispulum, Aphanes sp., Cerastium sp., Festuca rubra, Plantago coronopus, Thymus polytrichus,
less often Didymodon
acutus. On cliffs recorded with Fossombronia
'husnotii', Trichostomum
brachydontium, and very short
phanerogams.
A dioicous species for which capsules are unknown
in Britain.
