*1: Hayle Sands, 1861,
WC (PNZ) (Paton 1969a:
747).
*2: On steep bank of
sand at landward edge of dunes, 5 m alt., NW. of Rock, SW97,
2002, DTH 02-005 (BBSUK) (Rothero 2003:
60).
Most records are from calcareous sand in areas of
very short vegetation in dune grassland, including damp
hollows and dry sites, and sandy soil on partly bare ground
and short turf on unshaded tops of sea-cliffs. Associates
recorded: Barbula
convoluta, Bryum cf. algovicum, Homalothecium
lutescens, Hypnum
cupressiforme var.
lacunosum, Pleurochaete
squarrosa, Trichostomum
brachydontium, Festuca rubra, Holcus lanatus, Thymus polytrichus,
seedling Senecio
jacobaea; rarely Brachythecium
glareosum, Didymodon acutus. Also
recorded among very short vegetation on thin soil over ledge
of upper part of slaty crag near sea-cliffs; on steep sandy
bank at landward edge of dunes, partly shaded by Ash
sapling.
Occasionally c.fr. (three records, all vc1):
capsules immature 3; dehisced 10.
