*1: Trevaylor,Penzance, 1867, WC
(PNZ) (Paton 1969a:
739).
*2: St Breward, 1886,
HND (BM) (Paton 1969a:
739).
Rare in Isles of Scilly, where only record is of a
few stems found in Apr. 2003 growing among other mosses on a
damp shaded pathside bank at Higher Moor, St Mary's
(DTH).
Grows as scattered stems, or forming low lawns, but
the brown protonemata is often extensive where few
gametophores develop. Habitat notes from Cornwall are as
follows. Mostly on soil in wet places, usually in shade of
woodland or Grey Willow carr or scrub, e.g. flushes, stream,
ditch and river edges (extending low into flood-zones where it
sometimes forms large pure patches); small plants and
protonema also on tree bark (overhanging stream), rotting
branches, fern bases, rock and boulders. Bits on Molinia caerulea tussocks in
mire. Fern tussocks in swampy deciduous woodland. Flushes and
on unshaded streambank on and above sea-cliffs (with Bryum
pseudotriquetrum, Pellia epiphylla, Riccardia
chamedryfolia). Large plants on flushed steep slate rock
in quarry in deciduous woodland. Associates include Amblystegium serpens
var. serpens, Cephalozia
bicuspidata,
Kindbergia praelonga, Fissidens bryoides
var.
bryoides, Fissidens
bryoides var.
caespitans,
Hookeria lucens,
Hyocomium armoricum, Lophocolea
bidentata,
Lophocolea heterophylla, Mnium hornum, Pellia epiphylla, Pellia neesiana, Plagiomnium
undulatum,
Plagiothecium denticulatum var. denticulatum, Chrysosplenium
oppositifolium; less often Fissidens
polyphyllus,
Heterocladium heteropterum var. heteropterum, Trichostomum
tenuirostre.
Frequently c.fr.: capsules immature 1, 2, 9-12;
dehiscing 1, 2; dehisced 1-5.