*2: Crest of vertical
bank beside track in oakwood, NE. of Cabilla Manor, Warleggan
, 1988, JAP (BBSUK) (Blockeel 1989:
Not generally recognised in Britain until the
revision by Lewis & Smith (1978). Townsend (1995) revised
the nomenclature and showed that subspp. should not be
recognised (cf. Arts et
al. 1987, Smith 2004: 609).
Three records imply this species is tolerant of
heavy metals, or at least of zinc: Whitemoor: On low bank and
flat areas of china clay spoil near quarry, unshaded, close to
var. densifolia and
in and beside area where metal pipes and machine parts stored.
Hantergantick Quarry: patch among thick carpet of Bryum pallescens and
overlying steep edge of concrete at base of quarry building
made from galvanised iron. Greensplat: on china-clay spoil of
track beside quarry, close to galvanised shed (with Pohlia
Other habitat notes from Cornwall are as follows.
SW of Roche: on unshaded, sparsely vegetated, free-draining
low bank of china-clay spoil. SW. of Stenalees: in two places
on damp china clay spoil and silty soil of low bank near
china-clay works, unshaded to slightly shaded by scrub edge.
Goss Moor: on partly bare muddy soil exposed in acidic
grassland and on unshaded or partly bare edges of damp tracks
through disturbed ground near Grey Willow carrs (with Ceratodon purpureus, Cephaloziella
Only recorded with bulbils (short bulbils only at
Whitemoor and Goss Moor, long and short at Greensplat and
Roche). Not seen c.fr. (sporophytes are very rare in
to M.J. Wigginton in Hill et al. 1994: