Photo
by Dr M Lueth ©.
*1: Gulval nearPenzance, 1838, JR (TRU) (Paton 1969a: 742).
*2: Bodmin, before 1907, RVT (B) (Paton 1969a:
742).
Notes on habitats in C&S are as follows. Common
both as an epiphyte and on rock or masonry. As epiphyte,
commonest on Elder, frequent on Alder, Ash, Grey Willow,
Sycamore; few records from Apple, Blackthorn, Buddleja, Cortaderia, Cupressus macrocarpa,
elms, Gorse, hybrid poplars, oak, Traveller's Joy, White
Willow. Like other Orthotrichum,
epiphytic plants prefer bark of horizontal or inclined
branches or trunks to that on vertical ones, and they avoid
sites with heavy shade year-round (although tolerating
considerable shade in summer). Commoner overall than any of
its congeners, and often found in more exposed places than
those tolerated by O. affine or O. pulchellum (e.g. on
bushes on hillsides, and on bushes on tops of exposed
sea-cliffs). Frequent in areas of arable agriculture, even
occurring on Elders in hedgerows between arable fields where
high nutrient levels apparently exclude all other epiphytic
bryophytes. Also frequent on silted bark within flood-zone of
R. Tamar and large streams and recorded at upper edge of
inundation zone beside Stithians
Reservoir.
Also common on dry calcareous masonry (old and not
so old concrete and mortar), including walls, ruins, bridges,
graves, concrete fence-posts, a reservoir dam, masonry debris,
etc., where it grows on horizontal, vertical or inclined
surfaces, in unshaded or partly shaded places. Few records
also from serpentinite (natural boulders, on heath and by
pool), gabbro (natural boulder close to coast), a gravestone
and cliff bases of acidic rock, and several on granite
boulders or granite in walls. Unusual records also on old
tarmac of edge of tracks on coastal heaths (two records, both
cfr) and on thin hard soil and concrete dried onto metal of
farm roller parked at edge of field (many tufts,
cfr).
Associates when growing as an epiphyte are
essentially those listed as associates for O. affine; including Cololejeunea
minutissima, Cryphaea heteromalla,
Orthotrichum
tenellum, Syntrichia laevipila,
Ulota phyllantha,
Zygodon conoideus,
Zygodon
viridissimus var. viridissimus; also a
record with Syntrichia
papillosa. Associates when growing on concrete or mortar
are much like those listed for O. anomalum, including
Schistidium
crassipilum, Syntrichia montana, Syntrichia laevipila,
Tortula
muralis.
Foliar gemmae frequent, often present in abundance.
Commonly c.fr. [but differing from other Orthotrichum in that
young plants of this species were often recorded without
capsules]: capsules immature 1-12; dehiscing 1-11; dehisced
1-12.
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