*1: Near Chacewater,
1861, WC (NMW) (Paton 1969a:
698).
*2: Fowey, 1905, RWS
(TRU) (Paton 1969a:
698).
Notes on habitats in C&S are as follows. Soil
of bank beside old track in acid grassland. Clayey or firm
gritty soil of banks in woodland edges. Soil on paths, at
edges of tracks and on banks at edge of heath, near Grey
Willow carrs and in deciduous
and conifer woodlands, unshaded to moderately or sometimes
rather heavily shaded. Soil of tracks and banks in old
quarries, steep banks at track edges, peaty soil and firm
sediments of banks and flat ground at pool and reservoir edges
(including upper part of inundation-zone). Associates include
Archidium
alternifolium, Calypogeia arguta, Cephalozia
bicuspidata, Dicranella
heteromalla, Dicranella rufescens,
Diplophyllum
albicans, Nardia
scalaris, Riccardia
chamedryfolia, Scapania irrigua,
rarely Pogonatum
nanum. Damp, humic or clay soil of low banks or damp old
trackways on heaths (often growing from algal mat), of wet
peat banks in mires, flushed granitic rocks on sea-cliff (with
Blindia acuta, Scapania undulata) and
earthy banks in acidic flushes and beside heathland streams.
Once shallowly submerged in runnel in mesotrophic spring-fed
mire, with Scapania
undulata. Damp track in conifer
plantation.
Often dominant or forming extensive pure patches on
old copper-mine spoil, especially where copper-rich and damp,
and there often forming pure mats and locally dominant
(unshaded to lightly shaded), inland and on coasts; common
associates include Cephalozia
bicuspidata, Cephaloziella
divaricata, Cephaloziella
stellulifera, Ceratodon purpureus,
Dicranella varia,
Diplophyllum
albicans, Gymnocolea inflata, Pohlia annotina, Scapania compacta;
less common include Bryum alpinum, Calypogeia arguta, Pohlia andalusica,
rarely Ditrichum
plumbicola); often beneath open cover of phanerogams,
especially Calluna
vulgaris. Also on silty-clay, sandy silt and other
substrates on banks of streams draining old mine areas (where
it may be abundant and locally dominant), and in crevices of
walls built of old mine spoil, unshaded to part shaded.
Unshaded soil in areas of old china clay spoil and as colonist
on damp clay of banks, track edges, quarry floors, etc., in
old china clay quarries and near working quarries (sometimes
with Nardia
scalaris).
Perianths frequent: 1-4, 9-12. Frequently c.fr.:
immature 1-4, dehiscing 3-4, dehisced 3,
4.
