by Dr M Lueth ©.
*1: Madron Moor, 1883,
WC (MANCH) (Paton 1969a: 702). This record is
older than that listed by Castell (1950: 377) as new to
*2: Polperro, 1916, FR
(BM) (Paton 1969a:
Mainly identified when fertile or when non-fertile
material showed the characteristic leaf-stance (Paton 1999:
145), hence somewhat under-recorded.
Habitat notes from Cornwall are as follows. Thin
soil at edge of old gravelly track on mining ground, unshaded,
stellulifera, Ceratodon purpureus.
On sandy mine-spoil, unshaded. On unshaded metalliferous mine
spoil (associates include Campylopus
stellulifera, Ceratodon purpureus,
Thin soil on granite rocks in old quarries and on their spoil,
unshaded and lightly shaded, with Ditrichum
heteromallum, Nardia scalaris, Pogonatum urnigerum.
Soil and thin soil over serpentinite at edge of heath,
unshaded (often with Archidium
alternifolium). In and around active and old china clay
quarries as colonist on clayey banks, slopes and sometimes
flat areas with heath vegetation, also on crumbling or hard
granitic rocks or thin soil over rocks (often near Cephaloziella
purpureus, Marsupella sprucei, Nardia scalaris),
usually unshaded. Soil and thin soil on slope at base of
serpentinite crag and edges of paths etc. above serpentinite
and slaty sea-cliffs, unshaded (with Trichostomum
brachydontium, Weissia sp., often
intermixed with mosses). Acid soil on banks and slopes on and
above sea-cliffs and in unshaded, open areas of coastal heath
(close associates sometimes include Hypnum cupressiforme
Lophozia excisa, Polytrichum
juniperinum, Scleropodium touretii,
brachydontium, Conocephalum conicum).
Small mats on soil on little used old, stony track near gabbro
quarry; unshaded; soil on gravelly clearing in conifer wood,
Minions area and elsewhere: unshaded, thin soil and
edges of rocks in areas of copper-mine spoil. Among Ceratodon purpureus
introflexus on unshaded gravelly area of disused railway
track (on bridge top). Clayey ditch bank in clearing in
conifer plantation. Steep soil of part-shaded trackside bank.
Thin soil on ledge of old 'hedge' (with Scapania compacta).
Soil near rock on open heathland. With Campylopus flexuosus
on rock on low Cornish hedge. Among Campylopus flexuosus
on rocks in acid grassland slope. The common Cephaloziella on open
coastal heath in Isles of Scilly, often on open cliff tops or
coastal slopes, growing on peaty soil or mixed with mosses
such as Campylopus
introflexus. Almost unshaded firm sediment at top edge of
inundation-zone beside reservoir (with Calliergonella
Commonly with gemmae at stem tips: 1-4, 9, 11, 12.
Frequently cper: 1-6, 11, 12. Occasionally/frequently c.fr.:
immature 2, 3; dehiscing 1.