*1: Penzance, 1867, WC
(OXF) (Paton 1969a: 704).
*2: Helman Tor, 1906,
RWS (TRU) (Paton 1969a:
704).
Habitat notes from C&S are as follows. Usually
in patches on (often steep) surfaces of granitic rocks, less
often on thin soil over and at edges of the rocks, on rocky
hilltops, grassy hillsides, in old quarries, on banks of old
metalliferous mine spoil and on slope above sea-cliffs, on old
walls (especially north sides), most often unshaded, also part
to moderately shaded in deciduous woodlands. Locally abundant
on soil and ground litter amongst sparse Calluna vulgaris on
old mine area (with Hypnum jutlandicum).
One record on rock in area of china clay spoil. A few records
on bared peat or among other bryophytes on hummocks in open
mires. Also locally on Isles of Scilly on exposed peat of
N.-facing banks on coastal heaths. Unusual patch on very well
rotted remains of deciduous stump in young woodland near
stream, with mosses, part shaded. Sometimes also on acid soil
of woodland banks. Other associates include Diplophyllum albicans,
Lophozia
ventricosa, Plagiochila
asplenioides, less often Barbilophozia
attenuata, Bazzania
trilobata, Dicranum
scottianum, Marsupella emarginata
var. emarginata, Plagiochila
spinulosa.
Usually with foliar gemmae. One record c.fr.:
capsules dehisced 4.
