*1: Scilly, 1872, WC
(NMW) (Paton 1969a:
707).
*2: Fowey, 1905, RWS
(TRU) (Paton 1969a:
707).
Common as epiphyte on bark of trunks and branches
of shrubs and trees growing isolated, in groves or in scrub or
woodland. Recorded on Alder, Ash, Blackthorn (a locally
frequent phorophyte near coast), Grey Willow, Elder, elms,
Gorse, Hazel, Sessile Oak, Sycamore; single or few records
also from Apple, Broom, Buddleja, Cortaderia, Cotoneaster (tree and
low bushes), Garden Privet, Hawthorn, hybrid poplars, Picea abies (dead
twigs), Pinus
contorta, Rhododendron, Silver Birch, Sweet Chestnut,
White Willow, Wild Cherry, Wild Privet. Associates when
epiphyte include Cololejeunea
minutissima, Cryphaea heteromalla,
Frullania
tamarisci, Hypnum
cupressiforme var.
resupinatum, Metzgeria violacea, Metzgeria furcata, Metzgeria
consanguinea, Microlejeunea ulicina,
Neckera pumila, Orthotrichum affine,
Orthotrichum
diaphanum, Orthotrichum
pulchellum, Orthotrichum tenellum,
Radula complanata,
Ulota bruchii, Ulota crispa, Ulota phyllantha, Syntrichia laevipila,
Zygodon conoideus,
Zygodon viridissimus
var. viridissimus, rarely
Syntrichia
papillosa. Also often persisting for some years on fallen
branches. Frequently within flood-zone of River Tamar and
smaller streams; sometimes in upper part of inundation zone
beside reservoirs.
On rock outcrops, cliffs, boulders in quarry spoil
and rock in walls and 'hedges', gravestones (including
granitic rocks, gabbro, slates and serpentinite, and mortared
walls), usually unshaded to partly shaded, sometimes in
exposed places e.g. high on sea-cliffs. Associates on rocks
include Frullania
tamarisci, Grimmia
lisae, Hypnum
cupressiforme var.
resupinatum, Metzgeria furcata;
less often Cololejeunea
minutissima, Frullania microphylla
var.
microphylla.
Commonly c.per. (1-12) and frequently/commonly
c.fr.: capsules immature 1, 9-11, 12; dehiscing 11, 12;
dehisced 1-3, 7, 10-12.