Photo
by Dr M Lueth ©.
*1: Carn Brea, 1840, WC
(NMW) (Paton 1969a:
696).
*2: Helman Tor, Bodmin,
1906, RWS (TRU) (Paton 1969a:
696).
Habitat notes from C&S are as follows. Sides of
granitic rocks and boulders, thin and deeper soil (lithosols)
among rocks, and on banks in areas of old copper mine spoil;
unshaded or lightly shaded (once in small quantity part-shaded
in deciduous woodland); associates include Ceratodon purpureus,
Dicranum
scottianum, Didymodon insulanus,
Gymnocolea inflata,
Pseudocrossidium
hornschuchianum, Scapania compacta,
rarely Barbilophozia
attenuata. Soil on banks about old granite quarry.
Colonist on clay and gravelly substrates and granitic boulders
of banks around working china clay quarries, quarry floors and
on spoil heaps; usually unshaded, sometimes partly shaded.
Acid soil on banks, slopes and old tracks and thin soil over
granite and slate rocks high on sea-cliffs or above them
(sometimes in
exposed sites, as small fertile plants). Thin soil over
horizontal to vertical surfaces of granitic rocks at tors, and
on gravelly or gritty soil among granitic boulders on hilltop,
unshaded to part shaded (near Ceratodon purpureus,
Dicranum
scottianum, Polytrichum
juniperinum, Polytrichum piliferum,
Scapania gracilis).
A bit part shaded on soil over rock on wooded hillside. Thin
often humic soil on horizontal tops or steep or vertical sides
of old stone walls and 'hedges', with Diplophyllum albicans,
Scapania compacta.
Patch on steep rock in N.-facing wall beside lane. In Isles of
Scilly on peaty soil above cliffs and on coastal slopes, or
among rocks, often growing among mosses or other liverworts.
On damp china clay at unshaded edge of settling tank in works,
on thin layer of damp clay over horizontal concrete of ruined
works, unshaded (with Gymnocolea inflata, Scapania irrigua).
Damp peat of low bank beside acidic flush. Bodmin Moor: on
unshaded peat of low banks in and near mires. On healthy,
decaying or moribund sphagna and other vegetation and damp
bare peaty surfaces in mires, in flat areas or at bases of
hummocks (associates in mires include Calypogeia
muelleriana, Campylopus pyriformis,
Cephalozia
bicuspidata, Cephalozia connivens,
Kurzia pauciflora,
Kurzia sylvatica,
Kurzia
trichoclados, Lophozia incisa Mylia anomala, Odontoschisma sphagni,
Sphagnum
capillifolium; Drosera rotundifolia).
Usually (perhaps always?) with foliar gemmae, but
these sometimes sparse. Often c.per: 4, 11, ;occasionally
c.fr.: capsules immature 1, 10; mature 1, dehisced 10.
Abundantly fertile at Portheras, January 1998, over large
area