Range & Status
Mainly a Mediterranean species in Europe, reaching Crete and Albania in the east and extending north along Atlantic coasts to Scotland. Also occurs in Macaronesia and California. In Britain known in widely scattered localities in Wales and C. and S. England, with one in SW. Scotland (Kintyre).
Total of ca 20 localities known, on or near both coasts and inland. Doubtless underrecorded because of its small size and the critical nature of the genus.
Habitat & Ecology
Local and often occurring in small quantity, on crumbling slightly acidic, loamy soil in warm sheltered and lightly shaded places. Cornish records are from rocky crevices on cliffs, beside creeks, on rocky roadside cuttings, on stream banks and in a quarry. It is an autoicous species in which capsules are frequent, maturing in Cornwall from January to May. Foliar gemmae are common.
No detailed information, but likely to be at risk from increased shading as taller vegetation colonises its sites or from road works at some sites.