THE BRYOPHYTES OF CORNWALL
AND THE ISLES OF SCILLY


by
David T. Holyoak
     
Introduction -  Hornworts -  Liverworts -  Mosses -  Coastal influences -  Analysis -  Bibliography -  Acknowledgements -  Links -  CISFBR
A B C D F G H J K L M N O P R S T Layout notes Abbreviations People's initials Herbaria Full Text PDF

Liverworts

(MARCHANTIOPHYTA)
Adelanthus
decipiens


Anastrophyllum
minutum


Aneura
mirabilis


Aneura
pinguis


Aphanolejeunea
microscopica


Barbilophozia
attenuata


Barbilophozia
barbata


Barbilophozia
floerkei


Bazzania
trilobata


Blasia
pusilla


Blepharostoma
trichophyllum


Calypogeia
arguta


Calypogeia
fissa


Calypogeia
muelleriana


Calypogeia
sphagnicola


Cephalozia
bicuspidata


Cephalozia
catenulata


Cephalozia
connivens


Cephalozia
leucantha


Cephalozia
lunulifolia


Cephalozia
macrostachya


Cephalozia
macrostachya
var. macrostachya


Cephalozia
pleniceps


Cephaloziella
calyculata


Cephaloziella
dentata


Cephaloziella
divaricata


Cephaloziella
hampeana


Cephaloziella
integerrima


Cephaloziella
massalongi


Cephaloziella
massalongi s. l.


Cephaloziella
nicholsonii


Cephaloziella
rubella


Cephaloziella
spinigera


Cephaloziella
stellulifera


Cephaloziella
turneri


Chiloscyphus
pallescens


Chiloscyphus
polyanthos


Cladopodiella
fluitans


Cladopodiella
francisci


Cololejeunea
minutissima


Colura
calyptrifolia


Conocephalum
conicum s. l.


Conocephalum
conicum s. str.


Diplophyllum
albicans


Diplophyllum
obtusifolium


Douinia
ovata


Drepanolejeunea
hamatifolia


Dumortiera
hirsuta


Fossombronia
angulosa


Fossombronia
caespitiformis


Fossombronia
foveolata


Fossombronia
incurva


Fossombronia
loitlesbergeri


Fossombronia
maritima


Fossombronia
mittenii


Fossombronia
pusilla


Fossombronia
wondraczekii


Frullania
dilatata


Frullania
fragilifolia


Frullania
microphylla
var. microphylla


Frullania
tamarisci


Frullania
teneriffae


Gongylanthus
ericetorum


Gymnocolea
inflata


Gymnomitrion
concinnatum


Gymnomitrion
obtusum


Haplomitrium
hookeri


Harpalejeunea
molleri


Harpanthus
scutatus


Jamesoniella
autumnalis


Jamesoniella
undulifolia


Jubula
hutchinsiae


Jungermannia
atrovirens


Jungermannia
pumila


Kurzia
pauciflora


Kurzia
sylvatica


Kurzia
trichoclados


Leiocolea
badensis


Leiocolea
turbinata


Lejeunea
cavifolia


Lejeunea
lamacerina


Lejeunea
mandonii


Lejeunea
patens


Lepidozia
cupressina


Lepidozia
reptans


Lophocolea
bidentata


Lophocolea
bispinosa


Lophocolea
fragrans


Lophocolea
heterophylla


Lophocolea
semiteres


Lophozia
bicrenata


Lophozia
excisa


Lophozia
incisa


Lophozia
longiflora


Lophozia
sudetica


Lophozia
ventricosa


Lunularia
cruciata


Marchantia
polymorpha


Marchantia
polymorpha
subsp.
polymorpha


Marchantia
polymorpha
subsp.
ruderalis


Marchesinia
mackaii


Marsupella
emarginata
var. emarginata


Marsupella
funckii


Marsupella
profunda


Marsupella
sprucei


Metzgeria
conjugata


Metzgeria
consanguinea


Metzgeria
furcata


Metzgeria
violacea


Microlejeunea
ulicina


Moerckia
flotoviana


Mylia
anomala


Mylia
taylorii


Nardia
compressa


Nardia
geoscyphus


Nardia
scalaris


Nowellia
curvifolia


Odontoschisma
denudatum


Odontoschisma
sphagni


Pellia
endiviifolia


Pellia
epiphylla


Pellia
neesiana


Petalophyllum
ralfsii


Plagiochila
asplenioides


Plagiochila
bifaria


Plagiochila
exigua


Plagiochila
porelloides


Plagiochila
punctata


Plagiochila
spinulosa


Porella
arboris-vitae


Porella
cordaeana


Porella
obtusata


Porella
pinnata


Porella
platyphylla


Ptilidium
ciliare


Radula
complanata


Radula
lindenbergiana


Reboulia
hemisphaerica


Riccardia
chamedryfolia


Riccardia
incurvata


Riccardia
latifrons


Riccardia
multifida


Riccardia
palmata


Riccia
beyrichiana


Riccia
bifurca


Riccia
cavernosa


Riccia
ciliata


Riccia
crozalsii


Riccia
crystallina


Riccia
glauca


Riccia
huebeneriana


Riccia
michelii
var. ciliaris


Riccia
nigrella


Riccia
rhenana


Riccia
sorocarpa


Riccia
subbifurca


Saccogyna
viticulosa


Scapania
compacta


Scapania
curta


Scapania
gracilis


Scapania
irrigua


Scapania
nemorea


Scapania
paludicola


Scapania
scandica


Scapania
subalpina


Scapania
uliginosa


Scapania
umbrosa


Scapania
undulata


Solenostoma
caespiticium


Solenostoma
gracillimum


Solenostoma
hyalinum


Solenostoma
obovatum


Solenostoma
sphaerocarpum


Solenostoma
subellipticum


Southbya
tophacea


Sphaerocarpos
michelii


Sphaerocarpos
texanus


Targionia
hypophylla


Telaranea
europaea


Telaranea
murphyae


Trichocolea
tomentella


Tritomaria
exsectiformis


Tritomaria
quinquedentata


Riccia subbifurca Warnst. ex Croz. S12

(syn. R. warnstorfii auct. non. Limpr. ex Warnst.). Mediterranean-Atlantic Suboceanic element.


 


*1: Soil exposed on margin of Stithians Reservoir, 1973, JAP (BBSUK) (Corley 1979: 22,  Paton MS.: 8), as R. warnstorfii. [Housel Bay, Lizard, 1916, WEN, of Nicholson 1917 was redetermined as R. crozalsii by Nicholson 1918; see Paton 1969a: 691].

*2: Damp clay on edge of pool, Retew near Fraddon [SW95], 1962, JAP (BBSUK) (Paton 1969a: 691, as R. warnstorfii). [Record published as R. warnstorfii from track, Rocky Valley, Tintagel, JAP & JA, in Paton 1961: 150, is based on R. crozalsii, fide Paton 1969a: 691].

           

British records were revised by Paton (1990b) who showed that older records of R. warnstorfii belong here. It is now one of the commonest Riccia spp. in Cornwall and occasional in Isles of Scilly. R. subbifurca may have increased in the past two decades (change index +127%) but it seems more likely to have been under-recorded in the past, presumably because it was often overlooked as Riccia glauca.

 

Habitat notes from C&S are as follows. On damp soil of old track through woodland, partly or slightly shaded (with Anthoceros punctatus, Phaeoceros laevis, Riccia glauca, Riccia sorocarpa). On compressed soil of bare areas on and beside paths on sea-cliffs and slopes above them (over granitic, slate and serpentinite rocks); unshaded; with Archidium alternifolium, Bryum dichotomum, Fossombronia 'husnotii', Pleuridium acuminatum, Riccia sorocarpa, Scleropodium touretii, Trichostomum brachydontium, Weissia spp., Sedum anglicum; rarely Weissia longifolia var. longifolia. On rather loose mainly bare soil of ant-hill on slope above sea-cliff, unshaded. On partly bare patches of soil in old serpentinite quarry above sea-cliff, in trampled places and on banks, unshaded. Compressed, partly bare soil on little-used path at edge of heathland near coast (over serpentinite), unshaded. Kynance Cliff: plentiful in patches/rosettes on damp partly bare soil of small hollow on top of serpentinite sea-cliffs (with Archidium alternifolium, Trichostomum brachydontium, locally Riccia beyrichiana); path on slaty sea-cliff slope. Bit on soil of bare areas in pastures and near edge of grass-ley (with Bryum rubens, bit Riccia sorocarpa). Damp soil at edge of cauliflower field, unshaded (with Dicranella staphylina, Riccia glauca, Tortula truncata). Soil in flax, fallow, daffodil (and other horticultural) and cereal stubble fields (near Barbula convoluta, Bryum rubens, Dicranella schreberiana, Dicranella staphylina, Trichodon cylindricus, Ephemerum minutissimum, Phaeoceros laevis, Riccia glauca, Riccia sorocarpa, Tortula truncata, rarely Anthoceros agrestis, Fossombronia wondraczekii, Riccia crozalsii; in Isles of Scilly with Riccia crystallina, Spherocarpos spp.). Soil in arable (fallow) field (near Dicranella staphylina, Trichodon cylindricus, Riccia sorocarpa, Tortula truncata). Colonist on bare recently disturbed soil in unshaded clayey hollow (with Phaeoceros laevis, Pleuridium subulatum). Soil exposed on stream bank. Partly bare soil of old cattle-trodden tracks, unshaded (with Trichodon cylindricus, Ephemerum serratum, Ephemerum sessile, Pseudephemerum nitidum, Riccia glauca, Riccia sorocarpa, Tortula truncata). Usually unshaded or lightly shaded, one record on thin soil over gabbro rock partly shaded by Grey Willow scrub on Lowland Point (with Bryum bornholmense, Bryum rubens, Ephemerum serratum s. l., Fossombronia 'husnotii', Riccia beyrichiana). Sparsely on unshaded mud exposed in inundation zones beside Argal and Stithians Reservoirs (with Bryum klinggraeffii, Dicranella staphylina, Leptobryum pyriforme, Riccia sorocarpa).

 

Spores mature: immature 9, mature 2, 3, 7, 9, recently dehisced 9.