*1: On unshaded soil of low soil-heap, at edge of old
mine-spoil, Wheal Busy, E. of Scorrier, SW74, 1995, DTH 95-391
conf. TLB (BBSUK,
DTH) (Blockeel 1996: 45).
*2: On partly bare soil at edge of grassland in field
(car park) near coast, Talland Bay, SX225515, 1999, DTH 99-25
(Rothero 2010: 72).
For recognition of var. papillosum see Blockeel (1995). As
, this is a dubious taxon
because intermediates with var. cuspidatum are common.
Three good specimens exist from vc2 (DTH 99-25, 99-35,
00-829), but others (from SW84G and SW95M, both in DTH) show most but
not all characters of var. papillosum.
Notes on habitats in Cornwall are as follows. A
colonist of bare soil, of sandy, loamy or compacted textures
and circumneutral in reaction. Usually grows in open or
lightly shaded, less often in partly shaded sites (e.g. behind
north wall of church). Habitats typically include partly bare
patches beside paths and tracks, on banks, in disturbed areas
in grassland, sides of old wheel ruts, in arable fields
(stubble, flax stubble), in gardens, soil heaps, on top of
'hedge', in churchyards, on and above sea-cliffs, and on old
mining ground (including earthy mine-spoil). Also on dumped
soil of banks and flat areas near china clay quarries. Often
with or near var. cuspidatum, or
intermediate plants, and hence sharing list of more frequent
associates given for var. cuspidatum. Associates
recorded for this var. include Bryum dichotomum, B. rubens, Trichodon
Ephemerum minutissimum, Pleuridium
P. subulatum, Tortula
Commonly c.fr. (only recorded as this form when
spores mature): capsules immature 1-3, 11, 12; dehiscing 1-6,
9, 11, 12.