*1: Newlyn Cliff, 1861,
WC (OXF) (Paton 1969a:
753).
*2: St Minver, 1878,
RVT (B) (Paton 1969a:
753).
Habitat notes from C&S are as follows. Commonly
on circumneutral loam or friable soil, especially in humid and
sheltered places such as river banks, stream and ditch banks
in deciduous woodland or groves of trees, shady laneside
banks, where it may form extensive pure patches (or mixed with
Oxyrrhynchium
hians, Kindbergia
praelonga,
Fissidens taxifolius var. taxifolius, Plagiothecium
nemorale). Damp clay soil of bank under trees at edge of
pasture. Soil in churchyards, cemeteries, banks on and above
sea-cliffs. Sandy soil at base of granitic outcrop on slope
above coast, part shaded. Soil on sheltered and part shaded
'hedges' e.g. at wood edges, in gardens, in scrub above sea
cliff, by lane. Damp soil at edge of stream, part shaded.
Sheltered soil on slope above sea cliff. Patches low on
mortared walls, e.g. beside lane and above cliffs, part shaded
to rather heavily shaded. Also in hollow in ruin of wall. On
steep face of boulder of serpentinite in shade of scrub.
Extending onto slaty rocks in disused railway cutting, in
rather heavy shade. On rocks of shaded walls in humid sites,
also in small amount on unshaded laneside bank on soil above
wall. On masonry debris lying on old mining ground, only part
shaded. Soil among shaded ruined walls of old mine buildings
in woodland. Other associates recorded include Calypogeia arguta, Fissidens bryoides
var.
bryoides, Hookeria
lucens.
Occasionally c.fr.: capsules immature 1, 2, 11, 12;
dehiscing 2, [3], 11, 12; dehisced
3-5.