*1: Housel Bay, Lizard,
1917, WEN (Paton 1973a: 247, 248, 1974a: 162); specimens
collected from Housel Bay and Lizard Downs, 1916 to 1935, by
WEN, sometimes in the company of JBD, DAJ & HHK, are in
(BBSUK, BM, CGE, NMW, PC, S-PA) (Paton 1973a: 247).
*2: Soil on cliff top
near path, above S. side of Rocky Valley, between Boscastle
and Tintagel, SX08, 2000, JAP (BBSUK)
(Blackstock 2001: 36).
See Paton (1973a) for description and Paton (1994)
for elevation to species rank.
Although detailed microscopic study of mature
spores is usually necessary to identify this species,
well-grown plants lacking spores can be identified if they
combine purple rhizoids and a thick stem (although care needs
to be taken to avoid confusion with Fossombronia
Habitat notes from C&S are as follows. On Isles
of Scilly on coastal heaths, cliff tops, coastal slopes and in
short vegetation on acidic sand in a dune slack (Tresco),
often on partly bare trampled ground beside paths. Sometimes
forms large populations in partly bare damp peaty hollows in
vulgaris heaths. Recorded growing close to Fossombronia incurva.
A single atypical record from partly shaded loamy soil in a
garden ca 1 km
inland near Rocky Hill on St Mary's. On mainland on
compressed, partly bare, unshaded soil of track above
sea-cliffs, with Archidium
'husnotii', Lophozia excisa, Riccia crozalsii. Also
soil exposed on unshaded bank beside coastal
Only identified c.fr.(with mature spores, or when
grown-on). Capsules: immature 4, dehiscing