*1: Newlyn Cliff, 1864,
WC (PNZ) (Paton 1969a: 743). [An earlier
report: Penzance, AG, 1844, fide Paton MS.:
*2: Common, Trehane
near Probus, 1861, ES (TRU) (Paton 1969a:
Notes on its habitats in C&S are as follows.
Grows as slender stems appressed to bark when young, maturing
to form small patches, short wefts or mixtures with other
bryophytes, mainly as epiphyte on bark rich in nutrients and
bases. Commonest on Elder; also frequent to common on Ash,
Beech, Grey Willow, elms, Hazel, Ivy and Sycamore; other
records on Alder, oak spp. (two records one near coast, one
inland on Pedunculate Oak), old Gorse stems (three
localities); once each on Apple, Blackthorn, Buddleja, Cortaderia, Cotoneaster
Garden Privet, Hawthorn, Picea abies (once, on
dead twigs), Rowan, White Willow, Wild Privet. It occurs
mostly in sheltered sites with slight shade, avoiding places
exposed to severe desiccation. Although some sites are shaded
in summer by a deciduous leaf canopy it is scarce on heavily
shaded trees e.g. inside woodland.
It grows as a colonist on bare bark (vertical,
inclined or horizontal) or among other bryophytes, but cannot
compete with really luxuriant growth of pleurocarpous mosses
such as Hypnum
andoi and Isothecium myosuroides
myosuroides. Hence it occurs mainly at early or
intermediate stages of bryophyte colonisation of bark. On an
old Elder at Porkellis Moor young plants were established on
four-year old branches, it had begun to fruit on branches
whereas on 13-year old branches it had been almost
replaced by mats of Brachythecium
rutabulum and Kindbergia praelonga.
Frequent in scrub above sea-cliffs as well as
inland, occasionally even on Elders on upper parts of cliffs.
Often occurs within reach of the highest floods on riverside
trees and in inundation zones beside reservoirs, although
within flood-zone of R. Tamar it is generally found higher on
the trees than the rarer Dendrocryphaea
Common associates include Cololejeunea
Kindbergia praelonga, Frullania dilatata, Hypnum andoi, Hypnum cupressiforme
Orthotrichum affine, Orthotrichum
Orthotrichum pulchellum, Radula complanata, Ulota crispa, Ulota bruchii, Ulota phyllantha, Zygodon conoideus.
Others recorded include Brachythecium
Orthotrichum tenellum, Orthotrichum
Infrequently on rock or masonry, e.g. on granitic
boulders or posts (five records), rocks in walls (three
records), slaty boulders (once), serpentinite boulders (on
heath; in stream), concrete (of a river bridge and of
reservoir dam), sloping concrete in a pasture and dumped
asbestos-cement roofing sheets.
Commonly cfr; capsules immature 1-12; dehiscing 
7-12; dehisced [old 1-6] 7-12.