THE BRYOPHYTES OF CORNWALL
AND THE ISLES OF SCILLY


by
David T. Holyoak
     
Introduction -  Hornworts -  Liverworts -  Mosses -  Coastal influences -  Analysis -  Bibliography -  Acknowledgements -  Links -  CISFBR
A B C D F G H J K L M N O P R S T Layout notes Abbreviations People's initials Herbaria Full Text PDF

Liverworts

(MARCHANTIOPHYTA)
Adelanthus
decipiens


Anastrophyllum
minutum


Aneura
mirabilis


Aneura
pinguis


Aphanolejeunea
microscopica


Barbilophozia
attenuata


Barbilophozia
barbata


Barbilophozia
floerkei


Bazzania
trilobata


Blasia
pusilla


Blepharostoma
trichophyllum


Calypogeia
arguta


Calypogeia
fissa


Calypogeia
muelleriana


Calypogeia
sphagnicola


Cephalozia
bicuspidata


Cephalozia
catenulata


Cephalozia
connivens


Cephalozia
leucantha


Cephalozia
lunulifolia


Cephalozia
macrostachya


Cephalozia
macrostachya
var. macrostachya


Cephalozia
pleniceps


Cephaloziella
calyculata


Cephaloziella
dentata


Cephaloziella
divaricata


Cephaloziella
hampeana


Cephaloziella
integerrima


Cephaloziella
massalongi


Cephaloziella
massalongi s. l.


Cephaloziella
nicholsonii


Cephaloziella
rubella


Cephaloziella
spinigera


Cephaloziella
stellulifera


Cephaloziella
turneri


Chiloscyphus
pallescens


Chiloscyphus
polyanthos


Cladopodiella
fluitans


Cladopodiella
francisci


Cololejeunea
minutissima


Colura
calyptrifolia


Conocephalum
conicum s. l.


Conocephalum
conicum s. str.


Diplophyllum
albicans


Diplophyllum
obtusifolium


Douinia
ovata


Drepanolejeunea
hamatifolia


Dumortiera
hirsuta


Fossombronia
angulosa


Fossombronia
caespitiformis


Fossombronia
foveolata


Fossombronia
incurva


Fossombronia
loitlesbergeri


Fossombronia
maritima


Fossombronia
mittenii


Fossombronia
pusilla


Fossombronia
wondraczekii


Frullania
dilatata


Frullania
fragilifolia


Frullania
microphylla
var. microphylla


Frullania
tamarisci


Frullania
teneriffae


Gongylanthus
ericetorum


Gymnocolea
inflata


Gymnomitrion
concinnatum


Gymnomitrion
obtusum


Haplomitrium
hookeri


Harpalejeunea
molleri


Harpanthus
scutatus


Jamesoniella
autumnalis


Jamesoniella
undulifolia


Jubula
hutchinsiae


Jungermannia
atrovirens


Jungermannia
pumila


Kurzia
pauciflora


Kurzia
sylvatica


Kurzia
trichoclados


Leiocolea
badensis


Leiocolea
turbinata


Lejeunea
cavifolia


Lejeunea
lamacerina


Lejeunea
mandonii


Lejeunea
patens


Lepidozia
cupressina


Lepidozia
reptans


Lophocolea
bidentata


Lophocolea
bispinosa


Lophocolea
fragrans


Lophocolea
heterophylla


Lophocolea
semiteres


Lophozia
bicrenata


Lophozia
excisa


Lophozia
incisa


Lophozia
longiflora


Lophozia
sudetica


Lophozia
ventricosa


Lunularia
cruciata


Marchantia
polymorpha


Marchantia
polymorpha
subsp.
polymorpha


Marchantia
polymorpha
subsp.
ruderalis


Marchesinia
mackaii


Marsupella
emarginata
var. emarginata


Marsupella
funckii


Marsupella
profunda


Marsupella
sprucei


Metzgeria
conjugata


Metzgeria
consanguinea


Metzgeria
furcata


Metzgeria
violacea


Microlejeunea
ulicina


Moerckia
flotoviana


Mylia
anomala


Mylia
taylorii


Nardia
compressa


Nardia
geoscyphus


Nardia
scalaris


Nowellia
curvifolia


Odontoschisma
denudatum


Odontoschisma
sphagni


Pellia
endiviifolia


Pellia
epiphylla


Pellia
neesiana


Petalophyllum
ralfsii


Plagiochila
asplenioides


Plagiochila
bifaria


Plagiochila
exigua


Plagiochila
porelloides


Plagiochila
punctata


Plagiochila
spinulosa


Porella
arboris-vitae


Porella
cordaeana


Porella
obtusata


Porella
pinnata


Porella
platyphylla


Ptilidium
ciliare


Radula
complanata


Radula
lindenbergiana


Reboulia
hemisphaerica


Riccardia
chamedryfolia


Riccardia
incurvata


Riccardia
latifrons


Riccardia
multifida


Riccardia
palmata


Riccia
beyrichiana


Riccia
bifurca


Riccia
cavernosa


Riccia
ciliata


Riccia
crozalsii


Riccia
crystallina


Riccia
glauca


Riccia
huebeneriana


Riccia
michelii
var. ciliaris


Riccia
nigrella


Riccia
rhenana


Riccia
sorocarpa


Riccia
subbifurca


Saccogyna
viticulosa


Scapania
compacta


Scapania
curta


Scapania
gracilis


Scapania
irrigua


Scapania
nemorea


Scapania
paludicola


Scapania
scandica


Scapania
subalpina


Scapania
uliginosa


Scapania
umbrosa


Scapania
undulata


Solenostoma
caespiticium


Solenostoma
gracillimum


Solenostoma
hyalinum


Solenostoma
obovatum


Solenostoma
sphaerocarpum


Solenostoma
subellipticum


Southbya
tophacea


Sphaerocarpos
michelii


Sphaerocarpos
texanus


Targionia
hypophylla


Telaranea
europaea


Telaranea
murphyae


Trichocolea
tomentella


Tritomaria
exsectiformis


Tritomaria
quinquedentata


Cephaloziella integerrima (Lindb.) Warnst. NR:VU 12

Boreal-montane Suboceanic element.


 

 

 

*1: In rather bare areas over old, rocky mine-spoil, unshaded but sheltered by Gorse bushes, Lower Carnkie, SW64, 1996, DTH 96-257 (BBSUK, DTH) (Long 1997: 39-40).

*2: Slaty soil in quarry, E. of Withiel, Bodmin, 1963, JAP (BBSUK) (Corley 1983: 49).

           

The large majority of post-1950 British records are fromCornwall, including all sites currently known to hold the species. Easily overlooked in the field because it is small even for a Cephaloziella, and tends to grow as very short, open turfs as a colonist on small patches of bare substrates, rather than forming the procumbent mats of shoots typical of well-grown (and commonly fertile) material of the congeners that often grow nearby. Under the microscope the angular gemmae provide the first indication of its presence. Of the Cephaloziella recorded in similar habitats, only C. calyculata has similar gemmae, but that species is normally much larger in all vegetative parts and lacks any red or brown secondary pigementation, besides differing in characters of the perianth-mouth and bracts.

Recent records from ca 15 sites in vc1 and four in vc2 are all from old metalliferous mine sites, on mainly rather thin 'soil' over old mine-spoil that is at least sometimes copper-contaminated. Mainly recorded from unshaded places, less often from those lightly shaded (by Calluna vulgaris or by wall and Salix cinerea bushes, also once sheltered but not much shaded by tall Ulex europaeus). Once close to base of large galvanised-iron building. Twice on thin 'soil' in crevices of old walls retaining mine-spoil. Associated species include other small bryophytes of similar open habitats, among them Cephaloziella divaricata, Cephaloziella stellulifera, Ceratodon purpureus, Dicranella varia and Pohlia annotina, with Bryoerythrophyllum ferruginascens, Bryum dichotomum, Bryum pallens, Didymodon tophaceus, Ditrichum cornubicum, Ditrichum lineare and Conocephalum conicum at one site each. The copper-tolerant Cephaloziella massalongi, C. nicholsonii and Pohlia andalusica were also in similar habitats close by at several sites.

Found alone as small pure lawns and in small amounts in low 'turf' of Leptobarbula berica at Okeltor Mine near Calstock, on silty or sandy mine-spoil on top and ledges of low ruined walls at base of bank, an association that implies a calcareous substrate was present, but also present on flat ground nearby with acidophilous associates. Also on thin 'soil' over old concrete at Porkellis Moor, as well as on sandy substrates nearby.

Gemmae present on all gatherings examined and mainly common. Frequently cper, perianths present (but often rather few): 1, 9-11. Occasionally c.fr.: immature 3, dehiscing 3, dehisced 3.

Photomicrograph of gemmae.