THE BRYOPHYTES OF CORNWALL
AND THE ISLES OF SCILLY


by
David T. Holyoak
     
Introduction -  Hornworts -  Liverworts -  Mosses -  Coastal influences -  Analysis -  Bibliography -  Acknowledgements -  Links -  CISFBR
A B C D F G H J K L M N O P R S T Layout notes Abbreviations People's initials Herbaria Full Text PDF

Liverworts

(MARCHANTIOPHYTA)
Adelanthus
decipiens


Anastrophyllum
minutum


Aneura
mirabilis


Aneura
pinguis


Aphanolejeunea
microscopica


Barbilophozia
attenuata


Barbilophozia
barbata


Barbilophozia
floerkei


Bazzania
trilobata


Blasia
pusilla


Blepharostoma
trichophyllum


Calypogeia
arguta


Calypogeia
fissa


Calypogeia
muelleriana


Calypogeia
sphagnicola


Cephalozia
bicuspidata


Cephalozia
catenulata


Cephalozia
connivens


Cephalozia
leucantha


Cephalozia
lunulifolia


Cephalozia
macrostachya


Cephalozia
macrostachya
var. macrostachya


Cephalozia
pleniceps


Cephaloziella
calyculata


Cephaloziella
dentata


Cephaloziella
divaricata


Cephaloziella
hampeana


Cephaloziella
integerrima


Cephaloziella
massalongi


Cephaloziella
massalongi s. l.


Cephaloziella
nicholsonii


Cephaloziella
rubella


Cephaloziella
spinigera


Cephaloziella
stellulifera


Cephaloziella
turneri


Chiloscyphus
pallescens


Chiloscyphus
polyanthos


Cladopodiella
fluitans


Cladopodiella
francisci


Cololejeunea
minutissima


Colura
calyptrifolia


Conocephalum
conicum s. l.


Conocephalum
conicum s. str.


Diplophyllum
albicans


Diplophyllum
obtusifolium


Douinia
ovata


Drepanolejeunea
hamatifolia


Dumortiera
hirsuta


Fossombronia
angulosa


Fossombronia
caespitiformis


Fossombronia
foveolata


Fossombronia
incurva


Fossombronia
loitlesbergeri


Fossombronia
maritima


Fossombronia
mittenii


Fossombronia
pusilla


Fossombronia
wondraczekii


Frullania
dilatata


Frullania
fragilifolia


Frullania
microphylla
var. microphylla


Frullania
tamarisci


Frullania
teneriffae


Gongylanthus
ericetorum


Gymnocolea
inflata


Gymnomitrion
concinnatum


Gymnomitrion
obtusum


Haplomitrium
hookeri


Harpalejeunea
molleri


Harpanthus
scutatus


Jamesoniella
autumnalis


Jamesoniella
undulifolia


Jubula
hutchinsiae


Jungermannia
atrovirens


Jungermannia
pumila


Kurzia
pauciflora


Kurzia
sylvatica


Kurzia
trichoclados


Leiocolea
badensis


Leiocolea
turbinata


Lejeunea
cavifolia


Lejeunea
lamacerina


Lejeunea
mandonii


Lejeunea
patens


Lepidozia
cupressina


Lepidozia
reptans


Lophocolea
bidentata


Lophocolea
bispinosa


Lophocolea
fragrans


Lophocolea
heterophylla


Lophocolea
semiteres


Lophozia
bicrenata


Lophozia
excisa


Lophozia
incisa


Lophozia
longiflora


Lophozia
sudetica


Lophozia
ventricosa


Lunularia
cruciata


Marchantia
polymorpha


Marchantia
polymorpha
subsp.
polymorpha


Marchantia
polymorpha
subsp.
ruderalis


Marchesinia
mackaii


Marsupella
emarginata
var. emarginata


Marsupella
funckii


Marsupella
profunda


Marsupella
sprucei


Metzgeria
conjugata


Metzgeria
consanguinea


Metzgeria
furcata


Metzgeria
violacea


Microlejeunea
ulicina


Moerckia
flotoviana


Mylia
anomala


Mylia
taylorii


Nardia
compressa


Nardia
geoscyphus


Nardia
scalaris


Nowellia
curvifolia


Odontoschisma
denudatum


Odontoschisma
sphagni


Pellia
endiviifolia


Pellia
epiphylla


Pellia
neesiana


Petalophyllum
ralfsii


Plagiochila
asplenioides


Plagiochila
bifaria


Plagiochila
exigua


Plagiochila
porelloides


Plagiochila
punctata


Plagiochila
spinulosa


Porella
arboris-vitae


Porella
cordaeana


Porella
obtusata


Porella
pinnata


Porella
platyphylla


Ptilidium
ciliare


Radula
complanata


Radula
lindenbergiana


Reboulia
hemisphaerica


Riccardia
chamedryfolia


Riccardia
incurvata


Riccardia
latifrons


Riccardia
multifida


Riccardia
palmata


Riccia
beyrichiana


Riccia
bifurca


Riccia
cavernosa


Riccia
ciliata


Riccia
crozalsii


Riccia
crystallina


Riccia
glauca


Riccia
huebeneriana


Riccia
michelii
var. ciliaris


Riccia
nigrella


Riccia
rhenana


Riccia
sorocarpa


Riccia
subbifurca


Saccogyna
viticulosa


Scapania
compacta


Scapania
curta


Scapania
gracilis


Scapania
irrigua


Scapania
nemorea


Scapania
paludicola


Scapania
scandica


Scapania
subalpina


Scapania
uliginosa


Scapania
umbrosa


Scapania
undulata


Solenostoma
caespiticium


Solenostoma
gracillimum


Solenostoma
hyalinum


Solenostoma
obovatum


Solenostoma
sphaerocarpum


Solenostoma
subellipticum


Southbya
tophacea


Sphaerocarpos
michelii


Sphaerocarpos
texanus


Targionia
hypophylla


Telaranea
europaea


Telaranea
murphyae


Trichocolea
tomentella


Tritomaria
exsectiformis


Tritomaria
quinquedentata


Cephaloziella divaricata (Sm.) Schiffn. [S]12

(syn. C. starkei (Funck ex Nees) Schiffn. nom. illeg.; C. starkei var. scabra (M.Howe) L.Clark & Frye). Boreo-temperate Circumpolar element.


 

Photo by Dr M Lueth .

 

*1: Madron Moor, 1883, WC (MANCH) (Paton 1969a: 702). This record is older than that listed by Castell (1950: 377) as new to vc1.

*2: Polperro, 1916, FR (BM) (Paton 1969a: 702).

           

Mainly identified when fertile or when non-fertile material showed the characteristic leaf-stance (Paton 1999: 145), hence somewhat under-recorded.

Habitat notes from Cornwall are as follows. Thin soil at edge of old gravelly track on mining ground, unshaded, near Cephaloziella stellulifera, Ceratodon purpureus. On sandy mine-spoil, unshaded. On unshaded metalliferous mine spoil (associates include Campylopus introflexus, Cephaloziella stellulifera, Ceratodon purpureus, Lophozia bicrenata, Pohlia annotina). Thin soil on granite rocks in old quarries and on their spoil, unshaded and lightly shaded, with Ditrichum heteromallum, Nardia scalaris, Pogonatum urnigerum. Soil and thin soil over serpentinite at edge of heath, unshaded (often with Archidium alternifolium). In and around active and old china clay quarries as colonist on clayey banks, slopes and sometimes flat areas with heath vegetation, also on crumbling or hard granitic rocks or thin soil over rocks (often near Cephaloziella hampeana, Ceratodon purpureus, Marsupella sprucei, Nardia scalaris), usually unshaded. Soil and thin soil on slope at base of serpentinite crag and edges of paths etc. above serpentinite and slaty sea-cliffs, unshaded (with Trichostomum brachydontium, Weissia sp., often intermixed with mosses). Acid soil on banks and slopes on and above sea-cliffs and in unshaded, open areas of coastal heath (close associates sometimes include Hypnum cupressiforme var. lacunosum, Lophozia excisa, Polytrichum juniperinum, Scleropodium touretii, Tortula viridifolia, Trichostomum brachydontium, Conocephalum conicum). Small mats on soil on little used old, stony track near gabbro quarry; unshaded; soil on gravelly clearing in conifer wood, almost unshaded.

Minions area and elsewhere: unshaded, thin soil and edges of rocks in areas of copper-mine spoil. Among Ceratodon purpureus and Campylopus introflexus on unshaded gravelly area of disused railway track (on bridge top). Clayey ditch bank in clearing in conifer plantation. Steep soil of part-shaded trackside bank. Thin soil on ledge of old 'hedge' (with Scapania compacta). Soil near rock on open heathland. With Campylopus flexuosus on rock on low Cornish hedge. Among Campylopus flexuosus on rocks in acid grassland slope. The common Cephaloziella on open coastal heath in Isles of Scilly, often on open cliff tops or coastal slopes, growing on peaty soil or mixed with mosses such as Campylopus introflexus. Almost unshaded firm sediment at top edge of inundation-zone beside reservoir (with Calliergonella cuspidata, Fossombronia foveolata, Fossombronia incurva).

Commonly with gemmae at stem tips: 1-4, 9, 11, 12. Frequently cper: 1-6, 11, 12. Occasionally/frequently c.fr.: immature 2, 3; dehiscing 1.

Photomicrograph of two shoots, the right-hand with a female inflorescence.  Photo by Dr M Lueth .