*2: Steep damp gritty
soil of low bank at edge of river, unshaded, by De Lank River
just E. of Delford Bridge, ca 205 m alt., SX1175,
13 July 2006, DTH 06-231 (BBSUK, DTH)
(Holyoak 2007, Rothero 2008: 53).
The only British record of a species regarded as
Endangered in the Red
Data Book of European Bryophytes (Schumacker & Martiny
1995: 77). A few plants grew scattered in two small areas on
steep damp soil (humic and sandy) forming low banks at edge of
river (Holyoak 2007 & unpublished). Site is unshaded,
<1 m above (low) summer water level. Associated with Atrichum tenellum, Dicranella rufescens,
camptotrachela, immature Pellia epiphylla;
similar habitat close by also has Calypogeia arguta, Cephalozia
bicuspidata, Fissidens cf. bryoides var. caespitans, Philonotis sp., Pseudephemerum
nitidum. Phanerogams within <1m were: Agrostis capillaris,
decumbens, Euphrasia nemorosa, Juncus bulbosus, Plantago lanceolata,
In Portugal it also
grows on open soil, in sites subjected to grazing and
trampling by cattle, with 'an intermittent rivulet flowing
through acid grassland on peaty soil' (Sérgio et al.
Only specimen (collected 13 July 2006) has 4 good,
near mature capsules (see photo above).