THE BRYOPHYTES OF CORNWALL
AND THE ISLES OF SCILLY


by
David T. Holyoak
     
Introduction -  Hornworts -  Liverworts -  Mosses -  Coastal influences -  Analysis -  Bibliography -  Acknowledgements -  Links -  CISFBR
A B C D F G H J K L M N O P R S T Layout notes Abbreviations People's initials Herbaria Full Text PDF

Liverworts

(MARCHANTIOPHYTA)
Adelanthus
decipiens


Anastrophyllum
minutum


Aneura
mirabilis


Aneura
pinguis


Aphanolejeunea
microscopica


Barbilophozia
attenuata


Barbilophozia
barbata


Barbilophozia
floerkei


Bazzania
trilobata


Blasia
pusilla


Blepharostoma
trichophyllum


Calypogeia
arguta


Calypogeia
fissa


Calypogeia
muelleriana


Calypogeia
sphagnicola


Cephalozia
bicuspidata


Cephalozia
catenulata


Cephalozia
connivens


Cephalozia
leucantha


Cephalozia
lunulifolia


Cephalozia
macrostachya


Cephalozia
macrostachya
var. macrostachya


Cephalozia
pleniceps


Cephaloziella
calyculata


Cephaloziella
dentata


Cephaloziella
divaricata


Cephaloziella
hampeana


Cephaloziella
integerrima


Cephaloziella
massalongi


Cephaloziella
massalongi s. l.


Cephaloziella
nicholsonii


Cephaloziella
rubella


Cephaloziella
spinigera


Cephaloziella
stellulifera


Cephaloziella
turneri


Chiloscyphus
pallescens


Chiloscyphus
polyanthos


Cladopodiella
fluitans


Cladopodiella
francisci


Cololejeunea
minutissima


Colura
calyptrifolia


Conocephalum
conicum s. l.


Conocephalum
conicum s. str.


Diplophyllum
albicans


Diplophyllum
obtusifolium


Douinia
ovata


Drepanolejeunea
hamatifolia


Dumortiera
hirsuta


Fossombronia
angulosa


Fossombronia
caespitiformis


Fossombronia
foveolata


Fossombronia
incurva


Fossombronia
loitlesbergeri


Fossombronia
maritima


Fossombronia
mittenii


Fossombronia
pusilla


Fossombronia
wondraczekii


Frullania
dilatata


Frullania
fragilifolia


Frullania
microphylla
var. microphylla


Frullania
tamarisci


Frullania
teneriffae


Gongylanthus
ericetorum


Gymnocolea
inflata


Gymnomitrion
concinnatum


Gymnomitrion
obtusum


Haplomitrium
hookeri


Harpalejeunea
molleri


Harpanthus
scutatus


Jamesoniella
autumnalis


Jamesoniella
undulifolia


Jubula
hutchinsiae


Jungermannia
atrovirens


Jungermannia
pumila


Kurzia
pauciflora


Kurzia
sylvatica


Kurzia
trichoclados


Leiocolea
badensis


Leiocolea
turbinata


Lejeunea
cavifolia


Lejeunea
lamacerina


Lejeunea
mandonii


Lejeunea
patens


Lepidozia
cupressina


Lepidozia
reptans


Lophocolea
bidentata


Lophocolea
bispinosa


Lophocolea
fragrans


Lophocolea
heterophylla


Lophocolea
semiteres


Lophozia
bicrenata


Lophozia
excisa


Lophozia
incisa


Lophozia
longiflora


Lophozia
sudetica


Lophozia
ventricosa


Lunularia
cruciata


Marchantia
polymorpha


Marchantia
polymorpha
subsp.
polymorpha


Marchantia
polymorpha
subsp.
ruderalis


Marchesinia
mackaii


Marsupella
emarginata
var. emarginata


Marsupella
funckii


Marsupella
profunda


Marsupella
sprucei


Metzgeria
conjugata


Metzgeria
consanguinea


Metzgeria
furcata


Metzgeria
violacea


Microlejeunea
ulicina


Moerckia
flotoviana


Mylia
anomala


Mylia
taylorii


Nardia
compressa


Nardia
geoscyphus


Nardia
scalaris


Nowellia
curvifolia


Odontoschisma
denudatum


Odontoschisma
sphagni


Pellia
endiviifolia


Pellia
epiphylla


Pellia
neesiana


Petalophyllum
ralfsii


Plagiochila
asplenioides


Plagiochila
bifaria


Plagiochila
exigua


Plagiochila
porelloides


Plagiochila
punctata


Plagiochila
spinulosa


Porella
arboris-vitae


Porella
cordaeana


Porella
obtusata


Porella
pinnata


Porella
platyphylla


Ptilidium
ciliare


Radula
complanata


Radula
lindenbergiana


Reboulia
hemisphaerica


Riccardia
chamedryfolia


Riccardia
incurvata


Riccardia
latifrons


Riccardia
multifida


Riccardia
palmata


Riccia
beyrichiana


Riccia
bifurca


Riccia
cavernosa


Riccia
ciliata


Riccia
crozalsii


Riccia
crystallina


Riccia
glauca


Riccia
huebeneriana


Riccia
michelii
var. ciliaris


Riccia
nigrella


Riccia
rhenana


Riccia
sorocarpa


Riccia
subbifurca


Saccogyna
viticulosa


Scapania
compacta


Scapania
curta


Scapania
gracilis


Scapania
irrigua


Scapania
nemorea


Scapania
paludicola


Scapania
scandica


Scapania
subalpina


Scapania
uliginosa


Scapania
umbrosa


Scapania
undulata


Solenostoma
caespiticium


Solenostoma
gracillimum


Solenostoma
hyalinum


Solenostoma
obovatum


Solenostoma
sphaerocarpum


Solenostoma
subellipticum


Southbya
tophacea


Sphaerocarpos
michelii


Sphaerocarpos
texanus


Targionia
hypophylla


Telaranea
europaea


Telaranea
murphyae


Trichocolea
tomentella


Tritomaria
exsectiformis


Tritomaria
quinquedentata


Aneura pinguis (L.) Dumort. s. l. 12

(syn. Riccardia pinguis (L.) Gray). [A. pinguis s. l. is a Wide-boreal Circumpolar element.]


 


*1: Lamorna, 1882, WC (NMW) (Paton 1969a: 692).

*2: St Minver, 1906, RWS (TRU) (Paton 1969a: 692).

 

Recent molecular studies show that this taxon comprises a complex of cryptic species in Britain, with few morphological differences (D. G. Long pers. comm.).

           

Very large Aneura with thalli up to 10 mm wide from a shaded, flushed streambank near Lesnewth (DTH) raised hopes that A. maxima (Schiffn.) Steph. was at hand, but detailed study by JAP reidentified it as a large form of A. pinguis. Similarly large plants have also been seen on flushed ground with Chrysosplenium oppositifolium inside a Grey Willow carr near Trekeivesteps. Very large plants with long wide thalli were found a number of times in flushes in woodland (e.g. DTH 07-477), but although as large as A. maxima they do not show its other characters.

 

The wide range of habitat types occupied by this taxon may reflect occurrence of more than one morphologically cryptic species (see above). In Cornwall it occurs in at least three contrasting habitat types: (a) strongly acidic, peaty and unshaded; (b) mildly acidic, on mineral soils, often well shaded; (c) basic and unshaded.

 

Detailed habitat notes from Cornwall are as follows. On damp peaty or humic partly bare surfaces on heaths, on sphagna, mineral soils and damp humic litter in mires, stream edges and flushes above sea-cliffs (with Calliergonella cuspidata, Calypogeia fissa, Straminergon stramineum, Cephalozia bicuspidata, Kurzia pauciflora, Pellia endiviifolia, Riccardia chamedryfolia, Riccardia multifida, Sphagnum capillifolium, Sphagnum denticulatum; Anagallis tenella, Drosera rotundifolia, Narthecium ossifragum). On damp soil of partly bare patches along old tracks in heathland; bit in damp hollow on track in old mining ground, part shaded. Unusual record of small patches on open soil of heap amongst mine-spoil. Colonist on damp clay e.g. of flushed areas, of china clay quarry slopes, banks above working quarries and on spoil heaps, track edges, almost unshaded or partly shaded (with Anthoceros punctatus, Pellia neesiana, Philonotis fontana). Damp sand at edges of mica dams. On wet peaty humus of flush on slope above N.-facing sea-cliff, unshaded. With low mosses on unshaded, calcareous sandy substrate of flush on slope above sea-cliff (with Cratoneuron filicinum, Didymodon tophaceus). On open, flushed, calcareous sand of floor of sandpit in dunes (with Bryum pseudotriquetrum, Calliergonella cuspidata, Cratoneuron filicinum, Drepanocladus aduncus, Pellia endiviifolia, Petalophyllum ralfsii). Frequent in flushes and edges of small streams in deciduous woodland and Grey Willow carrs, growing on old fern bases, fern and other plant litter and intermixed with mosses, or on thin soil over wet rock, once on stream bank (associates include Calliergonella cuspidata, Chiloscyphus polyanthos s. l., Oxyrrhynchium speciosum, Pellia neesiana, Rhizomnium punctatum, Chrysosplenium oppositifolium). Beneath Grey Willows at upper edge of inundation-zone beside reservoir. On soil of gentle slope in open grove of deciduous trees. On wet clay near base of walls of ruin of china-clay dry, shaded by trees (very large plants). Small patch on partly bare soil on unshaded slope in old mine area, beside road. On wet ground amongst mine-spoil, at base of Juncus effusus. Damp soil of old metalliferous mine under edge of scrub (with Riccardia chamedryfolia). At normal water level on wet slaty rock in R. Camel with Hygroamblystegium fluviatile, partly shaded by trees and frequently inundated.

 

Commonly c.fr.: immature 1-3, 5, 12.